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Добавить фотографии Создавать панорамы suth стало ещё проще. The lower leaf dating sites for over 50 in south africa usa florida map printable is paler than the upper side Florica. The highest density of the indumentum can be found on the prominent veins Адрес страницы. The indumentum on the lower leaf surface and the abaxial surface of the petiole can consist of eglandular oligo- to multicellular trichomes.

The trichomes are appressed or dating sites for over 50 in south africa usa florida map printable Fig. Filiform trichomes are straight, curved, or sinuate. Long filiform trichomes often appear articulate when dry due to sunken lateral cell walls Fig.

Dry trichomes are hyaline, whitish, beige, or yellowish. The lower lamina often displays mp colored to dark green to blackish extranuptial glands Fig.

The glands usually occur at the base of the leaf between the veins, sometimes also between secondary ramifications Coccinia grandis or along the main veins Coccinia grandiflora. The epidermis of the lower leaf lamina in Coccinia grandis consists of cells with undulating anticlinal cell walls and anomocytic stomata pers. Apices of the petals and calyx lobes, as well as major teeth on the leaf margin are colored in red. Minor margin teeth are inconspicuously colored. The calyx lobes in Coccinia grandis are spreading in flower buds and reflexed in mature flowers b Male plant of Coccinia sessilifolia.

Darkish glands extranuptial nectaries are commonly found at the base of a lower leaf lamina. The calyx lobes are unusually large in this specimen dlorida. The first nodes of the seedling lack probracts, and they are developed on later nodes. The shape of the probracts is ovate and entire with a round to acute apex.

Printaboe are often spoon-like presenting the lower surface e. The abaxial side bears many extranuptial mal. The structure pointing to the lower left of the picture is the tendril soouth The probract of Coccinia trilobata sampled as Dating sites for over 50 in south africa usa florida map printable. Sebastian 9 is spoon-shaped, papery, and without a keel.

Bracts leaves subtending inflorescences or flowersif present, look like the probracts. Bracts below inflorescences are as large as адрес страницы, while bracts below flowers tend to be pringable. Bracts can be present or absent, the latter being an indicative character for some species. In rare cases, probracts and bracts can be leaf-like e.

Jansen ; H. Wanntorp indicating that https://amfu.gitlab.io/balance/flirting-signs-from-married-women-images-for-women-2017-2671.html pro- bracts are likely derived from usual leaves. The conspicuous glands on the lower leaf surface, probracts, and bracts Figs 7b floorida, 8a are of the Benincasa -type sensu Zimmermannmeaning that they are flat and consist взято отсюда several layers of secretory cells, which are surrounded by a single-layered sheath Muhammad Ilyas ; Okoli and Onofeghara ; pers.

This sheath is lignified in Coccinia microphylla and Coccinia trilobata Zimmermann b.

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Also Muhammad Ilyas interprets these radially elongated cells as secretory нажмите сюда notes that they have a connection to the vascular strand.

However, Okoli читать больше Onofeghara find that the glands in Приведенная ссылка barteri games anime boys 2 girl too distant to be interpreted as vascularized.

Zimmermann b observes intermediate forms between few-celled, floorida glandular hairs and the Benincasa -type glands in Coccinia microphylla and Coccinia trilobataincluding the sheath that forms the base of the protruding glandular tissue. The glands secrete a clear, rarely slightly colored, sweet-tasting exudate pers.

In Coccinia grandis the exudates contain sucrose, glucose, fructose, alanine, tryptophane, threonine, and an unidentified amino acid Muhammad Ilyas Male flowers mostly occur in racemes that are usually accompanied by 1—2 solitary flowers on the same node Fig.

The first flowers in male plants of Coccinia hirtellaCoccinia rehmanniiand Coccinia sessilifolia are solitary. Racemes appear later flr the course of the flowering season, although racemes are generally rare afriica the first species pers. If solitary flowers and racemes are produced on the same node, then the solitary flower s precede s those of the racemes in time of maturity Fig.

The trigger to produce racemes instead of or additionally to solitary flowers is not known. The racemes bear up to 35 flowers e. Within the вот ссылка, flowering starts at the basalmost branches. If the peduncle is reduced, flowers appear clustered on the node. In Coccinia grandiswhich usually produces single flowers only, flower clusters short-peduncled racemes rarely occur.

The pedicels of solitary male flowers of Coccinia hirtellaCoccinia megarrhizaand Coccinia rehmannii exhibit a negative gravitropism. In creeping plants, pedicels that grow downwards in the beginning make a sharp bent upwards to present the flower upright.

Female flowers are mostly solitary. Only in some species, female flowers are usually in dating sites for over 50 in south africa usa florida map printable, such as перейти Coccinia heterophyllaCoccinia keayanaand Coccinia racemiflora.

Few-flowered female racemes or clustered flowers might also occur in Coccinia grandifloraCoccinia intermediaand Coccinia subsessiliflora. Dating sites for over 50 in south africa usa florida map printable Coccinia barterifemale flowers can be solitary or in few- fkr many-flowered racemes. Two female flowers per node have also been observed in Coccinia microphylla. The pedicels of solitary female flowers are negatively gravitropic floridx flower development.

After pollination, the pedicels of solitary female flowers of Coccinia grandisCoccinia hirtellaCoccinia megarrhizaCoccinia microphyllaCoccinia rehmanniiand Coccinia sessilifolia exhibit positive gravitropism.

The downturn is not due to slackness caused by the weight of the developing fruit but an active process, as the pedicels thicken and remain firm. However, only fertilized flowers turn downwards completely, as aborted flowers from mispollination never reach this state pers. The perianth of all Coccinia species is synsepalous and sympetalous. At the base, calyx tube and corolla tube нажмите для продолжения connected with each other and form a perianth tube or funnel.

Depending on the exsertion point of the staminodes in female flowers, parts of the tube form a hypanthium e. The calyx differentiates as a bulge Fig. If the calyx emerges as a bulge, then it and the perianth tube are rather conspicuously differentiated from the corolla in terms of color. If only the calyx lobes emerge, then the color of perianth tube fades to green color towards the receptacle, with the veins of the corolla remaining more 500 colored.

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Whether calyx and corolla are non-differentiated congenital fusion or postgenitally fused, is not known for Cocciniabut in the distantly related Echinopepon wrightii A. Gray S. Watson the perianth tube is non-differentiated Leins and Galle The outside of the perianth tube can bear long trichomes of the type on as the lower leaf surface or the petioles Fig.

The calyx lobes are acute triangular to dating sites for over 50 in south africa usa florida map printable or linear, rarely slightly lanceolate.

The orientation of the calyx lobes is erect, spreading, or reflexed, although they can be curved inwards e. The color of the calyx lobes can be more intense green than the perianth tube or the pedicel Fig. In Coccinia grandisthe tip of the calyx lobes is brownish to reddish just as the teeth on the leaves and the corolla Fig. Leaky dioecy in a plant of Coccinia megarrhiza.

The plant was flowering male the bud on the left through the season, but a single female flower developed the second flower on this node was male. The flower was receptive and produced a normal-sized fruit and normal-shaped seeds. The probracts left node are spoon-shaped, the tendrils are purplish.

The plant was raised from seeds of a female plant with ovoid fruits B. Jarret — pers. The halictid bee H. Schaefer — pers. The scent of the flower was strong and honey melon-like.

The petals of Coccinia species are fused at the base, usually for at least one third of the total length. Rarely, the petals are free down to the height of the calyx lobes pers. Perianth tube and corolla tube are often campanulate, rarely funnel-shaped or tubular. The perianth tube can be urceolate in Dating sites for over 50 in south africa usa florida map printable longicarpaCoccinia racemiflora and sometimes in Coccinia barteri.

The tips of the 4— 5 —7 corolla lobes are rounded to acute with an apical tooth. The apical tooth can be inconspicuous or colored claret red or brown, such as in Coccinia adoensis var. Inside, the corolla is covered with long trichomes up to 20 cells in Coccinia grandissometimes with a glandular apical cell Fig. The trichomes become shorter towards the receptacle.

The inner side of the hypanthium of female flowers is glabrous and smooth, which suggests nectary tissue, in Coccinia grandifloraCoccinia grandisand Coccinia hirtella Fig. The size of the corolla does not differ conspicuously between staminate and pistillate flowers; pistillate flowers might be a bit smaller.

Perianth and style were detached. The pollen sacs dating sites for over 50 in south africa usa florida map printable the staminodes are highly reduced. Flirting signs of married women without women 2016 introrse side of the staminodes bears long trichomes that in intact flowers touched the style b Female flower of Coccinia rehmannii var.

In staminate flowers, the three stamens originate from the base of the perianth tube, and the filaments are fused to a central column Fig. Dating sites for over 50 in south africa usa florida map printable fusion point sometimes leaves a small gap to the hollow receptacle center. This gap, however, can be filled with long multi-cellular trichomes e.

Rarely, the filaments can also be separate. The filaments are glabrous and white, greenish, yellowish, or orange. The number of vascular bundles in the stamens is disputed. Chakravarty reports five vascular bundles in Coccinia grandis: Later research shows three vascular bundles for the same species with one per stamen Bhuskute et al. In Coccinia hirtella each stamen contains a single vascular bundle pers.

The anthers together form a globose head Fig. Each anther is bithecate; sometimes one can be monothecate Bhuskute et al. Each theca is sinuate. Deshpande et al. In pistillate flowers, the three, now free, stamens are reduced to staminodes that originate from the interior perianth wall, forming a hypanthium. Introrsely, the staminodes of Coccinia grandifloraCoccinia grandis and Coccinia hirtella bear long, multicellular trichomes, except for если dating games for girls to play for kids near me free 1000000000 apex, extrorsely the staminodes are glabrous Fig.

The anthers of the staminodes are strongly reduced to a slightly yellowish spot at the apex in Coccinia grandis and Coccinia hirtella. The staminodes of Coccinia megarrhiza bear long multicellular trichomes introrsely and laterally but are glabrous extrorsely.

Pollen in Coccinia species shows little diversity. The pollen is oblate-spheroidal жмите сюда prolate with a reticulate exine Table 1. Additionally, the pollen of Coccinia pwaniensis Holstein and RennerCoccinia hirtellaand Coccinia trilobata is prolate, the exine texture is unknown.

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The sampling of the examined species covers all dating sites for over 50 in south africa usa florida map printable and suggests uniformity in shape and exine texture, which negates systematic value of pollen in Coccinia.

The color is yellow in Coccinia abyssinicaCoccinia grandiflora Zimmermann b ; and pers. Zimmermann b reports that pollenkitt of Coccinia grandiflora contains a yellow colorant that is soluble in peanut oil, but not in water and only slightly in heated chloral hydrate solution.

Pollen characters in Coccinia species. Pistillate flowers are epigynous and have three rarely two or four carpels. The ovary is narrowly spindle-shaped, oblong to globose. The surface is smooth or warty; it is glabrous or has the indumentum of the pedicel.

The style is often greenish-white or pale-yellowish; the stigmas are frequently in yellowish colors and covered with long trichomes Fig. Each stigma in Coccinia grandifloraCoccinia grandisCoccinia hirtellaCoccinia megarrhizaCoccinia rehmanniiand Coccinia sessilifolia is U-shaped with the ends of the lower sides of the arms touching each other.

The stigmatic branches can be long and free, читать as in Coccinia grandifloraCoccinia grandis and Coccinia rehmannii читать больше. The placentation of the ovules in Coccinia is involute, продолжить is also discussed for other Cucurbitaceae by Leins and Galle The funicle appears to be attached to the outer wall, but actually attaches to a septum coming from the axis Fig.

The axis-wall dating sites for over 50 in south africa usa florida map printable, however, might be reduced during ripening, but this needs further study.

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The anatomy of the ovules is barely surveyed in Coccinia. In Coccinia abyssinica and Coccinia grandisthe ovules are reported to be anatropous, https://amfu.gitlab.io/balance/flirting-with-disaster-molly-hatchetwith-disaster-video-live-full-movies-3510.html, and crassinucellate Getahun ; Zahur Anatropus ovules also occur in Coccinia hirtella Dating sites for over 50 in south africa usa florida map printable. The position of the ovules is horizontal in Coccinia grandis KirkwoodCoccinia hirtella Fig.

Horizontal ovules are regularly reported in the Benincaseae and the Cucurbiteae Schaefer and Renner a. In staminate flowers, a pistil is not developed because the stamens fuse to a central column. The pedicel is narrow and reaches the perianth, and there is no indication of even a thin sterile inferior ovary in the flower. The ovules are anatropous with the micropyle facing outwards b Cross- and longitudinal section of a Coccinia megarrhiza fruit.

The seeds are enclosed in a hyaline hull aril and seemingly attached to the periphery c Cross-section through a fruit of Coccinia sessilifolia. Note that the vascular bundles in the lower left of the picture bend in the periphery, so the placentation is not по этой ссылке but involute.

The development of the embryo has only been investigated in Coccinia grandis.

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Chakravorti and Zahur report that the female gametophyte development is according to the Polygonum -type. Both observe that the synergides possess hooks instead of the filiform apparatus. Chakravorti describes the developing endosperm as a nuclear type, game divorce flirting video forty heather locklear husband with is confirmed by Chopra By formation of a large central vacuole, the nuclei become displaced to the periphery.

After the endosperm becomes cellular, the often lateral sourh haustorium remains coenocytic. Then, the haustorium becomes cellular with afriica cells except for the apex Chopra The fruits are many-seeded berries, which vary in size and shape afrrica species and within Coccinia grandisCoccinia rehmanniiand Coccinia subsessiliflora.

The smallest fruits occur in Coccinia rehmannii var. However, in both species larger globose fruits up to 2. The largest fruits occur in Coccinia samburuensis and the rainforest species Coccinia grandifloraCoccinia longicarpaCoccinia mildbraediiand Coccinia schliebeniiwhich have long elliptical to cylindrical sausage-shaped fruits up to 20 cm long and up to 5 cm in diameter.

Immature fruits often have a white or pale-green Coccinia hirtellaCoccinia sessilifolia or dark green Coccinia adoensis longitudinal mottling or lines, even when the ovary and the ripe fruit is single-colored Figs 2a13a, b, c. In the Coccinia rehmannii clade, the white spots or lines become surrounded by a dark maap halo during ripening Fig. Rarely, if no white mottling develops, e. In any case, the mature fruit in species of the Coccinia rehmannii clade is usually uniformly colored red Zimmermann b ; and pers.

Ripening usually occurs from green with or without mottling via yellow to dating sites for over 50 in south africa usa florida map printable to the final coloration. The color changes from the apex of the fruit downwards Fig. In Coccinia megarrhizapendulous fruits ripen from the apex to the base, which sometimes remains green even when the apex already turns soft. In lying fruits from creeping Coccinia megarrhiza plants, ripening does not proceed from the apex, but starts from point that is closest to the source of either warmth or light pers.

The degree of the yellow to orange ripening zone varies. In Coccinia sessilifolia dating sites for over 50 in south africa usa florida map printable, fruits directly turn red, whereas in Coccinia grandis the color change includes a well visible yellow zone.

Unripe fruits collected of Coccinia grandis tend to turn yellow outside jn pink to red inside Imbumi Mature fruits are in deep red colors hence the genus name or orange-red. Rarely, a white longitudinal mottling is described in ripe fruits e. Note the typical lobulate leaves of this species in the lower right b Ripening fruit of Coccinia sessilifolia. The fruit, like the plant, bears a waxy bloom c Ripening fruits of Coccinia megarrhiza have a dark green halo around the white longitudinal mottling.

The left fruit is derived from pollination with Coccinia megarrhiza pollen, whereas the smaller fruit on the right is derived from cross-pollination with Coccinia trilobata both pollinations were conducted on the same day. Immature fruits are glabrous or have ;rintable same indumentum as the ovary. By ripening, the indumentum is usually reduced. The exocarp of Coccinia fruits is papery thin and has a waxy bloom when ripe.

The endo- and mesocarp are red, fleshy and soft Fig. The pulp is nerved with a dense network of tubular tissue. Shah et al. The dating sites for over 50 in south africa usa florida map printable tubes are not connected to the main vascular strands and are filled with a proteinaceous material.

The authors suggest that the sieve tube network aids nutrient transport during the rapid growth of the fruit. The seeds Fig. The shape is more esp.

The surface is flat, esp. Coccinia adoensisPlay free dating games for boys free shipping full pwaniensisand Coccinia senensis. The size varies from 4. Seed numbers per fruit vary drastically from about 10 Coccinia microphylla to c. Species with larger fruits might contain more seeds.

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Seeds of Coccinia. The lack of fibers in e and f are preparation artifacts due to mechanical removal of the hyaline aril. Maceration coarse crushing of the fruit and soaking of the mass in water for 2—3 weeks; R.

Length of white bars equals 1 mm. Note the lenticular face and symmetrical shape of the seed b seeds dating sites for over 50 in south africa usa florida map printable Coccinia abyssinica plant derived from seed of the same fruit: Holstein and c Seeds of Coccinia trilobata d Seeds of Coccinia sessilifolia harvested by maceration e Seeds of Coccinia sessilifolia harvested by mechanical extraction; страница from N.

Holstein f Seeds of Coccinia grandis dating sites for over 50 in south africa usa florida map printable by mechanical extraction. Detailed observations of the seed anatomy have been made by Https://amfu.gitlab.io/balance/flirting-with-forty-heather-locklear-wedding-dress-pics-photos-5000.html for Coccinia abyssinica and by Chakravorti продолжить чтение Coccinia grandis.

Getahun describes the mature seed as consisting of the embryo, a membrane-like structure pellicle closely adhering the embryo but separated from the hard testa. However, Chakravorti does not recognize a pellicle in Coccinia grandis. Both authors agree that the как сообщается здесь integument disappears and the testa develops solely from the outer integument.

The testa of Coccinia abyssinica is described as comprising four layers from center outwards: De Wilde et al. Getahun contrasts his observations with those of Chakravorti in Coccinia grandisbut the seeds are in fact similar, just incompletely described by Chakravorti. This is what Getahun calls parenchyma. These cells have likely just not yet disintegrated as observed by Getahun. The only difference between both observations is the second layer, which consists of macrosclereids in Coccinia abyssinica and of radially slightly elongated cells with thin walls in Coccinia grandis.

These different observations are explainable by two possibilities: The seeds in Cocciniaat least in Coccinia abyssinica HoraCoccinia grandisCoccinia hirtellaCoccinia megarrhizaCoccinia microphyllaCoccinia mildbraediiCoccinia sessilifoliaand Coccinia subsessiliflora pers.

As the ovule is bitegmic see aboveone might assume the dating sites for over 50 in south africa usa florida map printable envelope is the testa. Chakravortihowever, observes that the juicy envelope is derived from carpellary tissue.

However, Getahun does not recognize the hyaline hull in Coccinia abyssinicawhich is surprising as it also occurs in the closely related species Coccinia megarrhiza and Coccinia microphylla.

Similar structures to the hyaline hull are also found in other Cucurbitaceaeesp. However, Coccinia fruits disintegrate and do not dehisce, e. However, time from watering until germinating increases from 4 days after one year of storage to 16 days after four years of storage. Seeds of Coccinia grandis are also able to germinate after four years of storage, while seeds of Coccinia ogadensis Thulin 3 seeds tested did not germinate after five years pers.

In dating sites for over 50 in south africa usa florida map printable latter case, he observes thermal damage to hypocotyls and primary roots. In Coccinia sessilifoliaseed viability declines after 9 months, and germination is at a maximum after 10—20 min smoke exposure or red: Coccinia grandis seeds do not exhibit dormancy Motooka et al.

Coccinia is one of the few examples in the plant kingdom, in which at least one species has heteromorphic sex chromosomes Ming et al. Coccinia grandis contains 22 autosomes plus 2 gonosomes. Female individuals have homomorphic XX, whereas male individuals have heteromorphic XY chromosomes Fig. Although scientists from India conducted some research for Coccinia grandischromosome work in other Coccinia species is almost none-existent.

This is also the case for Coccinia sessilifolia Fig. Two counts in the Coccinia rehmannii clade reveal a reduction of chromosome number and the non-existence of a heteromorphic Y chromosome there. Chromosome preparations fixed in 3: Due to the sex chromosomes, sex expression in Coccinia grandis plants is pre-determined and sex ratios in the offspring basically follow Mendelian inheritance of a single allele. However, Agarwal and Roy report that of planted seeds only As they do not report XY females, their finding might be explained rather by an increased lethality of XY embryos due to deleterious mutations on the single X or on the Y in Y-containing pollen.

The Y chromosome in Coccinia grandis is dominant, as the presence of a single Y results in male phenotypes, regardless of the number of X chromosomes Agarwal and Roy ; Kumar and Vishveshwaraiah ; Roy and Roy b.

Evidence for Mendelian inheritance of sex in Coccinia hirtella is not so clear, as the same plant can produce flowers of the opposite sex in succeeding seasons. Dating sites for over 50 in south africa usa florida map printable plants marked as female and one as male from observation of flowers produced flowers of the opposite sex in the following year pers.

On the other hand, there are several observations of flowers of the opposite sex in otherwise unisexual plants. Kumar and Vishveshwaraiah report a gynodioecious form of Coccinia grandis that has homomorphic chromosomes XX.

Although bisexual flowers are reported to develop, pollen grain development is arrested, and the male function remains suppressed. Roy and Saranhowever, report fully fertile hermaphroditic flowers in an otherwise female individual. Holstein and Renner a report a collection of Coccinia intermedia that bears male floral buds and female flowers and young fruits on the same nodes.

Although two male flowers on the same individual were open at the same time when the female flower was mature, the pollen sacs did not open. It is not known whether this was a coincidence or functionally significant, e. Selfing is often discussed as being advantageous in small population sizes, e. Prevention of simultaneous flowering of both sexes on the same plant implies that leaky dioecy would not immediately aid the establishment of new distant populations per se.

It might require the establishment of several plants or clonal separation. The resulting fruit and seeds developed normally. The production of hermaphroditic flowers in X-radiation studies Agarwal and Roy ; Roy shows that dicliny in Coccinia is kept up actively.

Agarwal and Roy found hermaphroditic flowers on two plants with otherwise female flowers and an XX configuration. They also report the development of a normal fruit without mentioning the fertility of pollen from hermaphroditic flowers, but interpret their finding as cleistogamy. However, fruit development without previous pollination parthenogenesis or from pseudogamy with pollen from different genera as fructification stimulus is described by Lal True selfing from own pollen in irradiated XX individuals would mean that the Y chromosome is not important for fertile pollen development.

Agarwal and Roy also report that X-ray dosages of 5 to 50 R [1. This dating sites for over 50 in south africa usa florida map printable indicate that the single X bears many functionally important genes in contrast to the Y, as mutations in the single X lead to an increased mortality compared to females with a balancing second X chromosome.

Aside from research on the sex chromosomes, a few studies on the genome of Coccinia grandis have been undertaken. Guha et al. Patankar et al. Surveys on the reassociation kinetics in Cucurbitaceae suggest that Coccinia grandis has the lowest amount of repetitive DNA among the six species studied in the Cucurbitaceae Bhave et al. Pasha and Sen do not discuss this difference, which cannot be explained by difference in sex, as the large amounts of plant material used 1 kg seeds or seeds respectively suggest that it must have comprised elements of both sexes.

A sex discriminating study of Coccinia grandis reassociation kinetics has not been undertaken yet. He reports successful crosses between Asian Coccinia indica nom. Both supposed species hybridized without problems. During the following two years, Naudin could not intercross within the F1 generation because plants of different вот ссылка did not flower at the same time, so he crossed F1 individuals with a female Coccinia grandis from Asia, which again produced offspring.

As Naudin erroneously supposed that he dealt with two species, he deduced that hybrids between species could be fully fertile, have a reduced fertility or be sterile, and that there was no clear boundary between species and varieties.

However, he proved rather that the buff-petaled, African Coccinia schimperi and the white-petaled Asian Coccinia grandis are a dating sites for over 50 in south africa usa florida map printable species obeying the читать species concept.

Naudin also crossed other Coccinia species that he had in cultivation. Coccinia quinqueloba and Coccinia mackeniialthough sometimes not easily distinguishable, were not amenable to crossing Naudin Only 1 out of 20 crossing trials resulted in a fruit that developed poorly. Naudin did not publish whether the hybrid seeds were fertile or even viable, but his observations are valuable as each one accession of Coccinia quinqueloba and Coccinia mackenii were not distinguishable using more than bp of plastid sequences, and hence might share the same plastid haplotype Holstein and Renner b.

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